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Research articles


  • Schulte to Bühne, H., Tobias, J.A., Durant, S.M., Pettorelli, N. (2020) Improving predictions of climate change–land use change interactions. Trends in Ecology & Evolution Online Early: PDF
  • Day, J.J., Martins, F.C., Tobias, J.A., Murrell, D.J. (2020) Contrasting trajectories of morphological diversification on continents and islands in the Afrotropical white-eye radiation. Journal of Biogeography Online Early PDF
  • Tobias, J.A., Ottenburghs, J., Pigot, A.L. (2020) Avian diversity: speciation, macroevolution and ecological function. Annual Reviews of Ecology, Evolution and Systematics 51: 533–60. PDF
  • Sheard, C., Neate-Clegg, M.H.C., Alioravainen, N., Jones, S.E.I., Vincent, C., MacGregor, H.E.A., Bregman, T.P., Claramunt, S., Tobias, J.A. (2020) Environmental and behavioural drivers of global gradients in dispersal inferred from avian wing morphology. Nature Communications 11: 2463. PDF
  • Sol, D., Trisos, C., Múrria, C., Jeliazkov, A., González-Lagos, C., Pigot, A.L., Ricotta, C., Swan, C.M., Tobias, J.A., Pavoine, S. (2020) The global impact of urbanisation on avian functional diversity. Ecology Letters 23: 962–972. PDF
  • Hawes, J.E., Viera, I.C.G., Magnago, L.F.S., Berenguer, E., Ferreira, J., Aragão, A., Cardoso, A., Lees, A.C., Lennox, G.D., Tobias, J.A., Waldron, A., Barlow, J. (2020) A large-scale assessment of plant dispersal traits across human-modified Amazonian forests. Journal of Ecology 108: 1373–1385. PDF
  • Jones, S.E.I., Tobias, J.A., Freeman, R., Portugal, S.J. (2020) Weak asymmetric interspecific aggression and divergent habitat preferences at an elevational contact zone between tropical songbirds. Ibis 162: 814–826. PDF
  • Hatfield, J.H., Orme, C.D.L., Lees, A.C., Tobias, J.A., Banks-Leite, C. (2020) Mediation of area and edge effects in forest fragments by adjacent land use. Conservation Biology 34: 395–404. PDF
  • Gallagher, R.V., Falster, D.S., Maitner, B., Salguero-Gómez , R., Vandvik, V., Pearse, W.D., Schneider, F.D., Kattge, J., Alroy, J., Ankenbrand, M., Andrew, S.C., Balk, M., Bland, L., Boyle, B., Bravo, C., Brennan, I., Carthey, A.J.R., Cavazos, B., Chown, S.L., Fadrique, B., Feng, X., Gibb, H., Halbritter, A.H., Hammock, J., Hogan, J.A., Holewa, H., Hope, M., Iversen, C.M., Jochum, M., Kearney, M., Keller, A., Mabee, P., Madin, J., Manning, P., McCormack, L., Michaletz, S.T., Park, D.S., Penone, C., Perez, T., Pineda-Munoz, S., Poelen, J., Ray, C.A., Rossetto, M., Sauquet, H., Sparrow, B., Spasojevic, M.J., Telford, R.J., Tobias, J.A., Violle, C., Walls, R., Weiss, K.C.B., Westoby, M., Wright, I.J., Enquist, B.J. (2020) Open Science principles for accelerating trait-based science across the Tree of Life. Nature Ecology & Evolution 4: 294–303. PDF
  • Pigot, A.L., Sheard, C., Miller, E.T., Bregman, T., Freeman, B., Roll, U., Seddon, N., Trisos, C.H., Weeks, B., Tobias, J.A. (2020) Macroevolutionary convergence connects morphological form to ecological function in birds. Nature Ecology & Evolution 4: 230–239. PDF
  • Kattge, J., Bönisch, G., Díaz, S., Lavorel, S., Prentice, I.C., et al. including Tobias, J.A. (2020) TRY plant trait database – enhanced coverage and open access. Global Change Biology 26: 189–190. PDF


  • Echeverri, A., Karp, D.S., Naidoo, R., Tobias, J.A., Zhao, J., Chan, K.M.A. (2019) Do avian functional traits predict cultural ecosystem services? People and Nature 2: 138–151. PDF
  • Habel, J.C., Tobias, J.A., Fischer, C. (2019) Movement of Afrotropical birds across a habitat mosaic: functional traits explain responses to land-use change better than species identity. Biotropica 51: 894–902. PDF
  • Betts, M.G., Wolf, C., Pfeifer, M., Banks-Leite, C., Arroyo-Rodríguez, V., Barlow, J., Bandini Ribeiro, D., Eigenbrod, F., Fletcher Jr., R.J., Hadley, A.S., Hawes, J.E., Holt, R.D., Klingbeil, B., Kormann, U., Lens, L., Levi, T., Mezger, D., Morante-Filho, J.C., Orme, D., Peres, C.A., Phalan, B.T., Pidgeon, A., Rangel, G.F.M., Ripple, W.J., Slade, E., Somarriba, E., Tobias, J.A., Tylianakis, J., Urbina-Cardona, J.N., Valente, J.J., Watling, J.I., Wells, K., Wearn, O.R., Wood, E., Young, R., Ewers, R.M. (2019) Extinction filters mediate the global effects of fragmentation. Science 366: 1236–1239. PDF
  • Freeman, B.G., Tobias, J.A., Schluter, D. (2019) Behavior influences range limits and patterns of coexistence across an elevational gradient in tropical bird diversity. Ecography 42: 1832–1840. PDF
  • Stoddard, M.C., Sheard, C., Yong, E.H., Akkaynak, D., Mahadevan, L., Tobias, J.A. (2019) Evolution of avian egg shape: underlying mechanisms and the importance of taxonomic scale. Ibis 161: 922–925. PDF
  • Santini, L., Butchart, S.H.M., Rondinini, C., Benítez-López, A., Hilbers, J.P., Schipper, A., Cengic, M., Tobias, J.A., Huijbregts, M.A.J. (2019) Applying habitat and population density models to land cover time series to inform IUCN Red List assessments. Conservation Biology 33: 1084–1092. PDF
  • Tobias, J.A., Pigot, A.L. (2019) Integrating behaviour and ecology into global biodiversity conservation strategies. Philosophical Transactions of the Royal Society B 374: 20190012. PDF
  • Cannon, P.G. Gilroy, J.J., Tobias, J.A., Medina Uribe, C.A., Haugaasen, T., Edwards, D.P. (2019) Land-sparing agriculture sustains higher levels of avian functional diversity. Global Change Biology 25: 1576–1590. PDF
  • Mayhew, R.J., Tobias, J.A., Bunnefeld, L., Dent, D.H. (2019) Connectivity with primary forest determines the value of regenerating tropical forests for bird conservation. Biotropica 51: 219–233. PDF
  • Kirschel, A.N.G., Seddon, N., Tobias, J.A. (2019) Range-wide spatial mapping reveals convergent character displacement of bird song. Proceedings of the Royal Society B 286: 20190443. PDF
  • Felice, R.N., Tobias, J.A., Pigot, A.L., Goswami, A. (2019) Dietary niche and the evolution of cranial morphology in birds. Proceedings of the Royal Society B 286: 20182677. PDF


  • Cooney, C.R., MacGregor, H.E.A., Seddon, N., Tobias, J.A. (2018) Multi-modal signalling in birds: re-examining a standard proxy for sexual selection. Proceedings of the Royal Society B 285: 20181557. Faculty of 1000 (special significance). PDF
  • Fecchio, A., Bell, J., Collins, M., Farias, I. Trisos, C., Tobias, J.A., Tkach, V., Weckstein, J., Ricklefs, R., Batalha-Filho, H. (2018) Diversification by host-switching and dispersal shaped the diversity and distribution of avian malaria parasites in Amazonia. Oikos 127: 1233–1242. PDF
  • Pigot, A., Jetz, W., Sheard, C., Tobias, J.A. (2018) The macroecological dynamics of species coexistence in birds. Nature Ecology & Evolution 2: 1012–1019. PDF
  • McEntee, J.P., Tobias, J.A., Sheard, C., Burleigh, J.G. (2018) Tempo and timing of ecological trait divergence in bird speciation. Nature Ecology & Evolution 2: 1120–1127. PDF
  • Chapman, P.M., Tobias, J.A., Edwards, D.P., Davies, R.G. (2018) Contrasting impacts of land-use change on phylogenetic and functional diversity of tropical forest birds. Journal of Applied Ecology 55: 1604–1614. PDF
  • Howard, C., Stephens, P.A., Tobias, J.A., Sheard, C., Butchart, S.H.M., Willis, S.G. (2018) Flight range, fuel load, and the impact of climate change on the journeys of migrant birds. Proceedings of the Royal Society B 285: 20172329. PDF
  • Derryberry, E., Seddon, N., Derryberry, G., Claramunt, S., Seeholzer, G., Brumfield, R.T., Tobias, J.A. (2018) Ecological drivers of song evolution in birds: disentangling the effects of habitat and morphology. Ecology & Evolution 8: 1890–1905. PDF
  • Bovo, A.A.A., Ferraz, K.M.P.M.B., Magioli, M., Alexandrino, E.R., Hasui, E., Ribeiro, M.C., Tobias, J.A. (2018) Habitat fragmentation narrows the distribution of avian functional traits associated with seed dispersal in tropical forest. Perspectives in Ecology & Conservation 16: 90–96. PDF
  • Drury, J.P., Tobias, J.A., Burns, K.J., Mason, N.A., Shultz, A.J., Morlon, H. (2018) Contrasting impacts of competition on ecological and social trait evolution in songbirds. PLoS Biology 16: e2003563. PDF
  • Hatfield, J.H., Orme, C.D.L., Tobias, J.A., Banks-Leite, C. (2018) Trait-based indicators of bird species sensitivity to habitat loss and fragmentation are effective within but not across datasets. Ecological Applications 28: 28–34. PDF


  • Waldron, A., Miller, D.C., Redding, D., Mooers, A., Kuhn, T.S., Nibbelink, N., Roberts, J.T., Tobias, J.A., Gittleman, J.L. (2017) Reductions in global biodiversity loss predicted from conservation spending. Nature 551: 364–367. PDF
  • Ulrich, W., Banks-Leite, C., De Coster, G., Habel, J.C., Matheve, H., Newmark, W.D., Tobias, J.A., Lens, L. (2017) Environmentally and behaviourally mediated co-occurrence of functional traits in bird communities of tropical forest fragments. Oikos 127: 274–284. PDF
  • Grether, G.F., Peiman, K.S., Tobias, J.A., Robinson, B.W. (2017) Causes and consequences of behavioral interference between species. Trends in Ecology & Evolution 32: 760–772. PDF
  • Thaxter, C.B., Buchanan, G.M., Carr, J., Butchart, S.H.M., Newbold, T., Green, R.E., Tobias, J.A., Foden, W.B., O’Brien, S., Pearce-Higgins, J.W. (2017) Bird and bat species’ global vulnerability to collision mortality at wind farms revealed through a trait-based assessment. Proceedings of the Royal Society B 284, 20170829. PDF
  • Stoddard, M.C., Yong, E.H., Akkaynak, D., Sheard, C., Tobias, J.A.*, Mahadevan, L.* (2017) Avian egg shape: form, function and evolution. Science 356: 1249–1254. (*equal contribution) (Altmetric = 1355) PDF
  • Cooney, C.R., Tobias, J.A., Weir, J.T., Botero, C.A., Seddon, N. (2017) Sexual selection, speciation, and constraints on geographical range overlap in birds. Ecology Letters 20: 863–871. PDF
  • Hosner, P., Tobias, J.A., Braun, E., Kimball, R. (2017) How do seemingly non-vagile clades accomplish trans-marine dispersal? Trait and dispersal evolution in the landfowl (Aves: Galliformes). Proceedings of the Royal Society B 284: 20170210. PDF
  • Bath, E., Bowden, S., Peters, C., Reddy, A., Tobias, J.A., Easton-Calabria, E., Seddon, N., Goodwin, S.F., Wigby, S. (2017) Sperm and sex peptide stimulate aggression in female Drosophila. Nature Ecology & Evolution 1: 0154. PDF
  • Fecchio, A., Svensson-Coelho, M., Bell, J., Ellis, V.A., Medeiros, M.C., Trisos, C.H., Blake, J.G., Loiselle, B.A., Tobias, J.A., Fanti, R., Coffey, E.D., De Faria, I.P., Pinho JB, Felix, G., Braga, E.M., Anciães, M., Tkach, V., Bates, J., Witt, C., Weckstein, J.D., Ricklefs, R.E., Farias, I.P. (2017) Host associations and turnover of haemosporidian parasites in manakins (Aves: Pipridae). Parasitology 144: 984–993. PDF
  • Mason, N.A., Burns, K.J., Tobias, J.A., Claramunt, S., Seddon, N., Derryberry, E.P. (2017) Song evolution, speciation, and vocal learning in passerine birds. Evolution 71: 786–796. PDF


  • Seddon, N., Mace, G., Naeem, S., Tobias, J.A., Pigot, A., Cavanagh, R., Mouillot, D., Vause, J., Walpole, M. (2016) Biodiversity in the Anthropocene: prospects and policy. Proceedings of the Royal Society B 283: 20162094. PDF
  • Bregman, T.P., Lees, A.C., MacGregor, H.E.A., Darski, B., de Moura, N.G., Aleixo, A., Barlow, J., Tobias, J.A. (2016) Using avian functional traits to quantify the impact of land-cover change on ecosystem processes linked to resilience in tropical forests. Proceedings of the Royal Society B 283: 20161289. PDF
  • Pigot, A., Bregman, T., Sheard, C., Daly, B., Etienne, R., Tobias, J.A. (2016) Quantifying species contributions to ecosystem processes: a global assessment of functional trait and phylogenetic metrics across avian seed-dispersal networks. Proceedings of the Royal Society B 283: 20161597. PDF
  • Ulrich, W., Lens, L., Tobias, J.A., Habel, J.C. (2016) Contrasting patterns in species richness and functional diversity in bird communities of East African cloud forest fragments. PLoS ONE 11: e0163338. PDF
  • Tobias, J.A., Sheard, C., Seddon, N., Meade, A., Cotton, A. J., Nakagawa, S. (2016) Territoriality, social bonds, and the evolution of communal signalling in birds. Frontiers in Ecology & Evolution 4: 74. PDF
  • Cooney, C.R., Seddon, N., Tobias, J.A. (2016) Widespread correlations between climatic niche evolution and species diversification in birds. Journal of Animal Ecology 85: 869–878. PDF
  • Collar, N.J., Lincoln D. C. Fishpool, L.D.C., del Hoyo, J., Pilgrim, J.D., Seddon, N., Spottiswoode, C.N., Tobias, J.A. (2016) Toward a scoring system for species delimitation: a response to Remsen. Journal of Field Ornithology 87: 104–115. PDF
  • Pigot, A., Tobias, J.A.*, Jetz, W.* (2016) Energetic constraints on species coexistence in birds. PLoS Biology 14: e1002407. (*equal contribution) PDF
  • Pigot, A., Trisos, C.H., Tobias, J.A. (2016) Functional traits reveal the expansion and packing of ecological niche space underlying an elevational diversity gradient in passerine birds. Proceedings of the Royal Society B 283: 20152013. PDF


  • Bregman, T.P., Lees, A.C., Seddon, N., MacGregor, H.E.A., Darski, B., Aleixo, A., Bonsall, M.B., Tobias, J.A. (2015) Species interactions regulate the collapse of biodiversity and ecosystem function in tropical forest fragments. Ecology 96: 2692–2704. PDF
  • Tobias, J.A. (2015) Biodiversity: Hidden impacts of logging. Nature 523: 163–164. PDF
  • Matthews, T.J., Sheard, C., Cottee‐Jones, H.E.W., Bregman, T.P., Tobias, J.A., Whittaker, R.J. (2015) Ecological traits reveal functional nestedness of bird communities in habitat islands: a global survey. Functional Ecology 124: 817–826. PDF
  • Bath, E., Wigby, S., Vincent, C., Tobias, J.A., Seddon, N. (2015) Condition, not eyespan, predicts contest outcome in female stalk-eyed flies, Teleopsis dalmanni. Ecology & Evolution 5: 1826–1836. PDF
  • Pigot, A., Tobias, J.A. (2015) Dispersal and the transition to sympatry in vertebrates. Proceedings of the Royal Society B 282: 20141929. PDF


  • Trisos, C.H., Petchey, O.L., Tobias, J.A. (2014) Unraveling the interplay of community assembly processes acting on multiple niche axes across spatial scales. American Naturalist 184: 593–608. PDF
  • Edwards, D.P., Tobias, J.A., Sheil, D., Meijaard, E., Laurance, W.F. (2014) Maintaining ecosystem function and services in logged tropical forests. Trends in Ecology & Evolution 29: 511–520. PDF
  • Tobias, J.A., Planque, R., Cram, D.L., Seddon, N. (2014) Species interactions and the structure of complex communication networks. Proceedings of the National Academy of Sciences USA 111: 1020–1025. PDF
  • Tobias, J.A., Cornwallis, C.K., Derryberry, E.P., Claramunt, S., Brumfield, R.T., Seddon, N. (2014) Species coexistence and the dynamics of phenotypic evolution in adaptive radiation. Nature 506: 359–363. PDF
  • Bregman, T.P., Sekercioglu, C.H., Tobias, J.A. (2014) Global patterns and predictors of bird species responses to forest fragmentation: Implications for ecosystem function and conservation. Biological Conservation 169: 372–383. PDF
  • Fayet, A., Tobias, J.A., Hintzen, R., Seddon, N. (2014) Immigration and dispersal are key determinants of cultural diversity in a songbird population. Behavioral Ecology 25: 744–753. PDF
  • Touchton, J.M., Seddon, N., Tobias, J.A. (2014) Captive rearing experiments confirm song development without learning in a tracheophone suboscine bird. PLoS ONE 9: e95746. PDF


  • Gonzalez, J.-C.T., Sheldon, B.C., Tobias, J.A. (2013) Environmental stability and the evolution of cooperative breeding in hornbills. Proceedings of the Royal Society B 280: 20131297. PDF
  • Seddon, N., Botero, C.A., Tobias, J.A., Dunn, P.O., MacGregor, H.E.A., Rubenstein, D.R., Uy, J.A.C., Weir, J.T., Whittingham, L.A., Safran, R.J. (2013) Sexual selection accelerates signal evolution during speciation in birds. Proceedings of the Royal Society B 280: 20131065. PDF
  • Gonzalez, J.-C.T., Sheldon, B.C., Collar, N.J., Tobias, J.A. (2013) A comprehensive molecular phylogeny for the hornbills (Aves: Bucerotidae). Molecular Phylogenetics and Evolution 67: 468–483. PDF
  • Pigot, A., Tobias, J.A. (2013). Species interactions constrain geographic range expansion over evolutionary time. Ecology Letters 63: 330–338. Faculty of 1000. PDF


  • Salisbury, C., Seddon, N., Cooney, C., Tobias, J.A. (2012) The latitudinal gradient in dispersal constraints: ecological specialisation drives diversification in tropical birds. Ecology Letters 15: 847–855. PDF
  • Tobias, J.A., Montgomerie, R., Lyon, B. (2012) The evolution of female ornaments and weaponry: sexual selection, social selection and ecological competition. Philosophical Transactions of the Royal Society B 367: 2274–2293. PDF
  • Tobias, J.A., Brawn, J.D., Brumfield, R.T., Derryberry, E.P., Kirschel, A.N.G., Seddon, N. (2012) The importance of suboscine birds as study systems in ecology and evolution. Ornitologia Neotropical 23: 259–272. PDF
  • Jankowski, J.E., Graham, C.H., Parra, J.L., Robinson, S.K., Seddon, N., Touchton, J.M., Tobias, J.A. (2012) The role of competition in structuring tropical bird communities. Ornitologia Neotropical 23: 115–124. PDF
  • Safran, R.J., Flaxman, S.M., Kopp, M., Irwin, D.E., Briggs, D., Evans, M.R., Funk, W.C., Gray, D.A., Hebet, E.A., Seddon, S., Scordato, E., Symes, L.B., Tobias, J.A., Toews, D.P.L., Uy, J.A.C. (2012) A robust new metric of phenotypic distance to estimate and compare multiple trait differences among populations. Current Zoology 58: 426–439. PDF
  • Derryberry E.P., Seddon N., Claramunt S., Tobias J.A., Baker A., Aleixo A., Brumfield, R.T. (2012) Correlated evolution of beak morphology and song in the Neotropical woodcreeper radiation. Evolution 66: 2784–2797. PDF


  • Tobias, J.A., Gamarra-Toledo, V., Garcia-Olaechea, D., Pulgarin, P.C. & Seddon, N. (2011) Year-round resource defence and the evolution of male and female song in suboscine birds: social armaments are mutual ornaments. Journal of Evolutionary Biology 24: 2118–2138. PDF
  • Barlow, J., Ewers, R.M., Anderson, L., Aragao, L., Baker, T., Boyd, E., Feldpausch, T., Gloor, M., Hall, T., Kouri, R., Malhi, Y., Milliken, W., Mulligan, M., Parry, L., Pennington, T., Peres, C.A., Phillips, O., Roman-Cuesta, R.M., Tobias, J.A., Gardner, T.A. (2011) Using learning networks to understand complex systems: a case study of biological, geophysical and social research in the Amazon. Biological Reviews 86: 457–474. PDF


  • Hoffmann, M. et al. (2010) The impact of conservation on the status of the world’s vertebrates. Science 330: 1503–1509. PDF
  • Tobias, J.A., Seddon, N., Spottiswoode, C. N., Pilgrim, J. D., Fishpool, L. D. C., Collar, N. J. (2010) Quantitative criteria for species delimitation. Ibis 152: 724–746. PDF
  • Tobias, J.A., Aben, J., Brumfield, R. T., Derryberry, E., Halfwerk, W., Slabbekoorn, H., Seddon, N. (2010) Song divergence by sensory drive in Amazonian birds. Evolution 64: 2820–2839. PDF
  • Seddon, N., Tobias, J.A. (2010) Character displacement from the receiver’s perspective: species and mate-recognition despite convergent signals in suboscine birds. Proceedings of the Royal Society B 277: 2475–2483. Faculty of 1000. PDF


  • Tobias, J.A., Seddon, N. (2009) Signal jamming mediates sexual conflict in a duetting bird. Current Biology 19: 577–582. PDF
  • Tobias, J.A., Seddon, N. (2009) Signal design and perception in Hypocnemis antbirds: evidence for convergent evolution via social selection. Evolution 63: 3169–3189. PDF
  • Seddon, N., Tobias, J. A., Eaton, M., Ödeen, A. (2009) Human vision can provide a valid proxy for avian perception of sexual dichromatism. Auk 127: 283–292. PDF
  • Tobias, J.A., Seddon, N. (2009) Sexual selection and ecological generalism are correlated in antbirds. Journal of Evolutionary Biology 22: 623–636. PDF


  • Seddon, N., Merrill, R. Tobias, J.A. (2008) Sexually selected traits predict patterns of species richness in a diverse clade of suboscine birds. American Naturalist 171: 620–631.  PDF
  • Tobias, J.A., Bates, J.M., Hackett, S.J., Seddon, N. (2008) Comment on: “The Latitudinal Gradient in Recent Speciation and Extinction Rates of Birds and Mammals”. Science 319: 901c. PDF
  • Tobias, J.A., Lebbin, D.J., Aleixo, A., Andersen, M.J., Guilherme, E., Hosner, P.A., Seddon, N. (2008) Distribution, behavior and conservation status of the Rufous Twistwing Cnipodectes superrufus. Wilson Journal of Ornithology 120: 38–49. PDF


  • Tobias, J.A., Brightsmith, D. (2007) Distribution, ecology and conservation status of the Blue-headed Macaw Primolius couloni. Biological Conservation 139: 126–138. PDF
  • Seddon, N., Tobias, J.A. (2007) Population size and habitat associations of the Long-tailed Ground-roller Uratelornis chimaera. Bird Conservation International 17: 1–13. PDF
  • Seddon, N., Tobias, J.A. (2007) Song divergence at the edge of Amazonia: an empirical test of the peripatric speciation model. Biological Journal of the Linnean Society 90: 173–188. PDF


  • Seddon, N., Tobias, J.A. (2006) Duets defend mates in a suboscine passerine, the warbling antbird (Hypocnemis cantator). Behavioral Ecology 17: 73–83. PDF


  • Seddon, N., Amos, W., Adcock, G.J., Johnson, P., Kraaijeveld, K., Kraaijeveld-Smit, F.J.L., Lee, W., Senapathi, G.D., Mulder, R.A., Tobias, J.A. (2005) Mating system, philopatry, and the role of kinship in the cooperatively breeding subdesert mesite Monias benschi. Molecular Ecology 14: 3573–3583. PDF


  • Tobias, J.A., Seddon, N. (2004) Breeding, foraging and vocal behavior in the White-throated Jacamar Brachygalba albogularis. Wilson Journal of Ornithology 115: 237–240. PDF
  • Seddon, N., Amos, W., Mulder, R., Tobias, J.A. (2004) Male heterozygosity predicts territory size, song structure and reproductive success in a cooperatively breeding bird. Proceedings of the Royal Society B 271: 1823–1829. PDF


  • Seddon, N., Tobias, J.A. (2003) Communal singing in the cooperatively breeding subdesert mesite Monias benschi: evidence for numerical assessment? Journal of Avian Biology 34: 72–80. PDF
  • Seddon, N., Tobias, J.A., Butchart, S.H.M. (2003) Group living, territoriality and breeding behaviour in the Subdesert Mesite Monias benschi. Ibis 145: 277–294. PDF
  • Tobias, J.A., Seddon, N. (2003) Vocalization and display in the Long-tailed Ground-roller Uratelornis chimaera. Wilson Journal of Ornithology 115: 193–196. PDF


  • Seddon, N., Tobias, J.A., Alvarez, A. (2002) Vocal communication in the Pale-winged Trumpeter Psophia leucoptera: repertoire, context and functional reference. Behaviour 139: 1331–1359 .PDF
  • Tobias, J.A., Seddon, N. (2002) Estimating population size in the Subdesert Mesite: new methods and implications for conservation. Biological Conservation 108: 199–212. PDF
  • Tobias, J.A., Seddon, N. (2002) Female begging in European robins: do neighbours eavesdrop for extra-pair copulations? Behavioral Ecology 13: 637–642. PDF
  • Ekstrom, J.M.M., Jones, J.P.G., Willis, J., Tobias, J.A., Dutson, G., Barré, N. (2002) New information on the distribution, status and conservation of terrestrial bird species in New Caledonia. Emu 102: 197–207.  PDF


  • Clay, R.P., López Lanús, B., Tobias, J.A., Lowen, J.C., Mazar Barnett, J. (2000) The display of the White-winged Nightjar Caprimulgus candicans. Journal of Field Ornithology 71: 619–626. PDF
  • Mazar Barnett, M., Kirwan, G.M., Pearman, M., Naka, L.N., Tobias, J.A. (2000) Rediscovery and subsequent observations of Kaempfer’s Tody-tyrant Hemitriccus kaempferi in Santa Catarina, Brazil, with notes on conservation, life history and plumage. Bird Conservation International 10: 371–379. PDF
  • Seddon, N., Tobias, J.A., Yount, J.W., Ramanampamonjy, J.R., Butchart, S.H.M., Randrianizahana, H. (2000) Conservation issues and priorities in the Mikea Forest of south-west Madagascar. Oryx 34: 287–304. PDF
  • Tobias, J.A., Seddon, N. (2000) Territoriality as a paternity guard in the European robin Erithacus rubecula. Animal Behaviour 60: 165–173.  PDF


  • Brooks, T., Tobias, J.A., Balmford, A. (1999) Deforestation and bird extinctions in the Atlantic Forest. Animal Conservation 2: 211–222. PDF


  • Dabelsteen, T., McGregor, P.K., Holland, J., Tobias, J.A., Pedersen, S.B. (1997) The signal function of overlapping singing in male robins. Animal Behaviour 53: 249–256. PDF
  • Tobias, J.A. (1997) Asymmetric territorial contests in the European robin: the role of settlement costs. Animal Behaviour 54: 9–21. PDF
  • Tobias, J.A. (1997) Food availability as a determinant of pairing behaviour in the European robin. Journal of Animal Ecology 66: 629–639. PDF


  • Tobias, J.A., Williams, R.S.R. (1996) Notes on the behaviour of the Masked Saltator in southern Ecuador. Auk 113: 942–944. PDF

Books and chapters

  • Tobias, J.A, Şekercioğlu, Ç.H., Vargas, F.H. (2013) Bird conservation in tropical ecosystems: challenges and opportunities. In Key Topics in Conservation Biology, vol. 2. eds D. MacDonald, K. Willis, pp. 258–276. John Wiley & Sons, London. PDF
  • Tobias, J.A., Bennun, L., Stattersfield, A. (2005) Listening to the birds. Pp. 169–177 in Wildlife Conservation Society (2005) State of the wild 2006: a global portrait of wildlife, wildlands, and oceans. Island Press, Washington, DC. PDF
  • Fishpool, J., Tobias, J.A. (2005) Family Pycnonotidae (Bulbuls). Pp. 124–250 in del Hoyo et al. (eds) Handbook of the Birds of the World. Volume 10. Lynx Edicions, Barcelona.
  • Tobias, J.A. (2002) Family Galbulidae (Jacamars). Pp. 74–101 in del Hoyo et al. (eds) Handbook of the Birds of the World, Volume 7. Lynx Edicions, Barcelona.
  • Collar, N., Andreev, A.V., Chan, S., Crosby, M.J., Subramanya S., Tobias, J.A. (2001) Threatened birds of Asia. BirdLife International, Cambridge, UK. [3038 pp.]

Guides and reports

  • Tobias, J.A. (2009) Antbirds of Peru and Bolivia. The Field Museum, Chicago. (Photoguide) PDF
  • Tobias, J.A., Koch, P., Merkord, C. (2008) Hummingbirds of the Madre de Dios watershed. The Field Museum, Chicago. (Photoguide) PDF
  • BirdLife International (2003) Saving Asia’s Threatened Birds: a guide for government and civil society. BirdLife International, Cambridge, UK. [100 pp.]
  • Tobias, J.A. (2003) Rio Tinto Biodiversity Strategy: conservation opportunities at Rio Tinto operations worldwide. BirdLife International and World Conservation Monitoring Centre, Cambridge, UK.
  • Tobias, J.A. (2003) Status of the Blue-throated Macaw Ara glaucogularis in Beni department, Bolivia. Asociación Armonia/Loro Parque, Santa Cruz, Bolivia. [50 pp.]
  • Isherwood, I.S., Willis, J.D.A., Edwards, T.R.K., Ekstrom, J.M.M., Kuriake, S., Lubis, I.R., Notanubun, H., Putnarubun, J., Robinson-Dean, J.C., Tobias, J.A. (1997) Biological surveys and conservation priorities in Seram, Indonesia. CSB Conservation Publications, Cambridge, UK. [160 pp.]
  • Lowen, J.C., Bartrina, T.M., Clay, R.P., Tobias, J.A. (1996) Biological surveys and conservation priorities in eastern Paraguay. CSB Conservation Publications, Cambridge, UK. [200 pp.]
  • Tobias, J.A. (1997) Nakai-Nam Theun Social and Environmental Action Plan. Wildlife Conservation Society/IUCN, Vientiane, Lao PDR. [400 pp.]
  • Williams, R.S.R., Tobias, J.A. (1994) The conservation of southern Ecuador’s threatened avifauna. BirdLife International, Cambridge, UK. [150 pp.]

Specialist articles

  • Echeverri, A., Karp, D., Tobias, J.A. (2020) Loved or loathed: what makes some Neotropical birds more popular than others? Neotropical Birding 26: 15–21. PDF
  • Lowen, J.C., Lees, A.C., Tobias, J.A. (2019) Frontiers of knowledge: a quarter-century of Neotropical discovery. Neotropical Birding 25: 11–30. PDF
  • Tobias, J.A., Seddon, N. (2007) Nine bird species new to Bolivia and notes on other significant records. Bulletin of the British Ornithologists’ Club 127: 49–84. PDF
  • Tobias, J.A., Seddon, N. (2007) Ornithological notes from southern Bolivia. Bulletin of the British Ornithologists’ Club 127: 293–300. PDF
  • Tobias, J.A., (2007) Range extensions and recent additions to the Bolivian avifauna. Neotropical Birding 2: 36–39.
  • Tobias, J.A., Butchart, S.H.M., Collar, N.J. (2006) Lost and found: a gap analysis for the Neotropical avifauna. Neotropical Birding 1: 4–22. PDF
  • Tobias, J.A., del Hoyo, J. (2006) Birding Bolivia: putting two rare curassows on the map. Neotropical Birding 1: 60–65.
  • Butchart, S.H.M., Collar, N.J., Crosby, M.J., Tobias, J.A. (2005) “Lost” and poorly known birds: top targets for Asian conservation. BirdingASIA 3: 41–49. PDF
  • Tobias, J.A. (2003) Notes on breeding behaviour in Black-faced Cotinga Conioptilon mcilhennyi. Cotinga 19: 80–81.
  • Tobias, J.A., Seddon, N. (2002) Reading the sand: identifying bird tracks in Madagascar’s spiny forest. African Bird Club Bulletin 9: 12–15.
  • Parr, J.W.K., Benstead, P.J., Tobias, J.A. (2002) A first nest record for the Fruithunter Chlamydochaera jefferyi. Forktail 18: 154–155. PDF
  • Tobias, J.A., Ekstrom, J.M.M. (2002) The New Caledonian Owlet-nightjar Aegotheles savesi rediscovered? Bulletin of the British Ornithologists’ Club 122: 282–283. PDF
  • Tobias, J.A., Seddon, N. (2002) First nest record for Scarlet-shouldered Parrotlet. Cotinga 18: 104.
  • Naka, L.N., Mazar Barnett, J., Kirwan, G.M., Tobias, J.A., De Azevedo, M.A.G. (2000) New and noteworthy state records from Santa Catarina state, Brazil. Bulletin of the British Ornithologists’ Club 120: 237–250. PDF
  • Tobias, J.A. (2000) Critically Threatened: White-eyed River-martin. Oriental Bird Club Bulletin 31: 45–48. PDF
  • Clay, R.P, Tobias, J.A., Lowen, J.C., Beadle, D. (1998) Field identification of Phylloscartes and Phyllomyias tyrannulets in the Atlantic Forest region. Cotinga 10: 82–95. PDF
  • Ekstrom, J.M.M., Tobias, J.A., Robinson-Dean, J. (1998) A survey of Lore Lindu National Park, Sulawesi. Oriental Bird Club Bulletin 28: 37–39.
  • Isherwood, I.S., Edwards, T.R.K., Ekstrom, J.M.M., Robinson-Dean, J.C., Tobias, J.A., Willis, J.D.A. (1998) New information on the birds of Seram, Indonesia, and their conservation. Oriental Bird Club Bulletin 28: 18–21.
  • Tobias, J.A., Davidson, P., Robichaud, W. (1998) Nakai-Nam Theun: can development save one of South-East Asia’s last wildernesses? Oriental Bird Club Bulletin 28: 24–29.
  • Lowen, J.C., Clay, R.P., Mazar Barnett, J., Madroño-N., A., Pearman, M., López Lanús, B., Tobias, J.A., Liley, D.C., Brooks, T.M., Esquivel, E.Z., Reid, J.M. (1997) New and noteworthy observations on the Paraguayan avifauna. Bulletin of the British Ornithologists’ Club 117: 275–293. PDF
  • Lowen, J.C., Bartrina, L., Brooks, T.M., Clay, R.P., Tobias, J.A. (1996) Project Yacutinga ’95: bird surveys and conservation priorities in Paraguay. Cotinga 5: 14–19. PDF
  • Tobias, J.A., Williams, R.S.R. (1996) Threatened Formicivora antwrens of Rio de Janeiro state, Brazil. Cotinga 5: 62–66. PDF